(海岸性甲虫)

Nishikawa, M. (2010) A colluvial slope as a hibernating habitat of littoral beetles in eastern Hokkaido, north Japan. Elytra, Tokyo, 38(1): 61-64.
Asano, M., S. Inokuchi and H. Kojima (2010) Records of some marine beetles new to the fauna of Sado Island, central Japan. Elytra, Tokyo, 38(1): 71-72.