Kudo, Y., H. Kojima, H. Yoshitake and Y. G. Baba (2012) Population structure of the flightless supralittoral weevil, Aphela gotoi (Coleoptera, Curculionidae) inferred from mitochondrial COI sequence. Jpn. J. Syst. Ent., 18(2): 195-202.／ハマベゾウムシの分子系統．日本海側と太平洋側で大きく分かれる．
