A. E. Marvaldi et al. (2009) Structural alignment of 18S and 28S rDNA sequences provides insights into phylogeny of Phytophaga (Coleoptera: Curculionoidea and Chrysomeloidea) . Zoologica Scripta, 38, 1, January 2009, pp63–77.／ハムシ上科の単系統性が支持されている．
A. E. Marvaldi et al. (2009) Structural alignment of 18S and 28S rDNA sequences provides insights into phylogeny of Phytophaga (Coleoptera: Curculionoidea and Chrysomeloidea) . Zoologica Scripta, 38, 1, January 2009, pp63–77.／ハムシ上科の単系統性が支持されている．